Scientific Articles
2026
Gomez Doris, Arias Monica, Dhennin Marina, Aubier Thomas G, Andraud Christine, Barla Pascal, Elias Marianne. Transparency, the ultimate camoufage? Background dependency and influence of background lightness. Proceedings B 2075 (1): 2026310. https://doi.org/10.1098/rspb.2026.0310
Gomez Doris, Charline Pinna, Violaine Ossola, Christine Andraud, Serge Berthier, Stephan Borensztajn, Marianne Elias. Optical and mechanical constraints drive the evolution of wing membrane thickness in clearwing butterflies. Journal of Evolutionary Biology.
2025
Ossola Violaine, Fabien Pottier, Charline Pinna, Katia Bougiouri, Aurélie Tournié, Anne Michelin, Christine Andraud, Doris Gomez et Marianne Elias. It is cool to be clear: Transparency induces a thermal cost in clearwing butterflies. Functional Ecology 39, n°9 (2025): 2452-2464 https://doi.org/10.1111/1365-2435.70110
2021
Arias Monica, Lucie Leroy, Clement Madec, Louane Matos, Cynthia Tedore, Marianne Elias, et Doris Gomez. « Partial Wing Transparency Works Better When Disrupting Wing Edges: Evidence from a Field Experiment ». Journal of Evolutionary Biology 34, no 11 (2021): 1840‑46. https://doi.org/10.1111/JEB.13943.
Gomez Doris, Charline Pinna, Jonathan Pairraire, Monica Arias, Jerome Barbut, Aaron Pomerantz, Willy Daney de Marcillac, et al. « Wing Transparency in Butterflies and Moths: Structural Diversity, Optical Properties and Ecological Relevance ». Ecological Monographs 91, no 4 (2021): e01475. https://doi.org/10.1002/ecm.1475.
Pinna Charline, Maëlle Vilbert, Stephan Borenztajn, Willy Daney de Marcillac, Florence Piron-Prunier, Aaron Pomerantz, Nipam Patel, et al. « Convergence in Light Transmission Properties of Transparent Wing Areas in Clearwing Mimetic Butterflies ». ELife 10 (2021): e69080. https://doi.org/10.7554/eLife.69080.
Pomerantz Aaron F., Radwanul H. Siddique, Elizabeth I. Cash, Yuriko Kishi, Charline Pinna, Kasia Hammar, Doris Gomez, Marianne Elias, et Nipam H. Patel. « Developmental, Cellular, and Biochemical Basis of Transparency in the Glasswing Butterfly Greta Oto ». Journal of Experimental Biology accepted, no 224 (2021): eb237917. https://doi.org/10.1242/jeb.237917.
2020
Arias Monica, Marianne Elias, Christine Andraud, Serge Berthier, et Doris Gomez. « Transparency Improves Concealment in Cryptically Coloured Moths ». Journal of Evolutionary Biology 33, no 2 (2020): 247‑52. https://doi.org/10.1111/jeb.13560.
2019
Arias Monica, Johanna Mappes, Charlotte Desbois, Swanne Gordon, Melanie McClure, Marianne Elias, Ossi Nokelainen, et Doris Gomez. « Transparency Reduces Predator Detection in Mimetic Clearwing Butterflies ». Functional Ecology 33, no 6 (2019): 33(6), 1110-1119. https://doi.org/10.1111/1365-2435.13315.
McClure Melanie, Corentin Clerc, Charlotte Desbois, Aimilia Meichanetzoglou, Marion Cau, Lucie Bastin-Héline, Javier Bacigalupo, et al. « Why has transparency evolved in aposematic butterflies? Insights from the largest radiation of aposematic butterflies, the Ithomiini ». Proceedings of the Royal Society B: Biological Sciences 286, no 1901 (24 avril 2019): 20182769. https://doi.org/10.1098/rspb.2018.2769.
Preprints
Arias Mónica, Jérôme Barbut, Rodolphe Rougerie, Manon Dutry, Mireia Kohler, Baptiste Laulan, Caroline Paillard, et al. « Ambient Light and Mimicry as Drivers of Wing Transparency in Lepidoptera ». BioRxiv, 2020, 2020.06.26.172932. https://doi.org/10.1101/2020.06.26.172932v3.
Gomez Doris, Jonathan Pairraire, Charline Pinna, Monica Arias, Celine Houssin, Jerome Barbut, Serge Berthier, Christine Andraud, Thierry Ondarcuhu, et Marianne Elias. « Hydrophobicity in Clearwing Lepidoptera: Impact of Scale Micro and Nanostructure, and Trade-off with Optical Transparency », BioRxiv 2021. https://doi.org/10.1101/2021.10.11.463886.
PhD dissertations
Violaine Ossola. Thermal adaptation in tropical clearwing butterflies. PhD Dissertation, Université Paris Cité. 2023. here
Charline Pinna. Evolution of wing transparency in mimetic Lepidoptera: optical properties, structural modifications and selection for convergence. PhD Dissertation, MNHN Paris. 2021. here
Press releases and dissemination articles
generalist journals
article in Le Monde on our work on transparent butterflies
specialised science websites
article (in French) on the CNRS platform on our work
Cairn info website (in French) on science about our work on transparent butterflies
technoscience info website (in French) on our work on transparent butterflies
in English For some butterflies, reason for wing adaptation is clear
techno